Date of birth	1953
Date of death	2008
Occupation	professori
Field of Activity	eläintiede biometria ekologia
Identifiers	0000000116164198 (isni) http://urn.fi/URN:NBN:fi:au:finaf:000056426 (finaf)
Sources	Helsingin Sanomat 21.9.2008
Associated Groups	Helsingin yliopisto (-)